The skin of the desert lark protects from water loss via a ceramide-rich lipid ratio.

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"Adjustments of lipid ratios to favor ceramides over free fatty acids and sterols have also been shown to correlate with reductions of TEWL [transepidermal water loss] in desert larks (Haugen et al., 2003a,b). The comparatively higher ratios of ceramides in stratum corneum allow the lipid lamellae of the permeability barrier to exist in a more highly ordered crystalline phase, which creates a tighter barrier to water vapor diffusion (Velkova and Lafleur, 2002; Bouwstra et al., 2003b)." (Lillywhite 2006:218)

Journal article
Water relations of tetrapod integumentJournal of Experimental BiologyJanuary 4, 2006
H. B. Lillywhite

Journal article
Phenotypic flexibility in cutaneous water loss and lipids of the stratum corneumJournal of Experimental BiologySeptember 9, 2003
M. J. Haugen

Journal article
Lipids of the Stratum Corneum Vary with Cutaneous Water Loss among Larks along a Temperature‐Moisture GradientPhysiological and Biochemical ZoologyMarch 3, 2004
Michael Haugen, Joseph B. Williams, Philip Wertz, B. Irene Tieleman

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