Special glands in male sea lamprey gills release a bile that acts as a sex pheromone to attract long-distance females.

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“We show that reproductively mature male sea lampreys release a bile acid that acts as a potent sex pheromone, inducing preference and searching behavior in ovulated female lampreys..the male of this fish species signals both its reproductive status and location to females by secreting a pheromone that can act over long distance.” (Li et al. 2002:138)

Journal article
Bile acid secreted by male sea lamprey that acts as a sex phermoneScience Vol. 296, Issue 5565, pp. 138-141April 5, 2002
Li, W; Scott, A.P.; Siefkes, M.J.; Yan, H.; Liu, Q.; Yun, S.; Gage, D.

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